This was the second Coupe built and the first completed at Carrozzeria Gransport. It competed in nine FIA races (LeMans, Reims, Goodwood Tourist Trophy, Tour de France, Daytona, Sebring, Oulton Park TT, LeMans, Enna), won four FIA events (LeMans '64, Tourist Trophy '64, Daytona '65, Sebring '65) and one event during the Tour de France (Rouen). The car was driven through this period by Dan Gurney, Bob Bondurant, Maurice Trintignant, Bernard de St. Auban, Jo Schlesser, Hal Keck, Jack Sears and Dick Thompson. At LeMans in 1964, drivers Gurney and Bondurant, clocking over 315 KPH on the Mulsanne straight, took First Place in the GT III Class.

CSX2299 was painted Viking Blue with two white stripes in 1964 and repainted Guardsman Blue with two larger longitudinal white stripes and a transverse white stripe across the nose in 1965. In 1964 this car set lap records at Le Mans, Reims and Rouen and the race distance record at Le Mans and Goodwood. In 1965 the car set the lap record at Oulton Park.

CSX2299 was the last Daytona coupe sold by Shelby with a Bill of Sale which read "the number one Cobra Daytona coupe". It is currently owned by Larry H. Miller and resides at the Miller Motorsports Park museum in Tooele, Utah.